The Rise of Muscovy
When the Mongols invaded the lands of Kievan Rus', Moscow was an insignificant trading outpost in the principality of Vladimir-Suzdal'. The outpost's remote, forested location offered some security from Mongol attack and occupation, and a number of rivers provided access to the Baltic and Black seas and to the Caucasus region. More important to Moscow's development in what became the state of Muscovy, however, was its rule by a series of princes who were ambitious, determined, and lucky. The first ruler of the principality of Muscovy, Daniil Aleksandrovich (d. 1303), secured the principality for his branch of the Rurik Dynasty. His son, Ivan I (r. 1325-40), known as Ivan Kalita ("Money Bags"), obtained the title "Grand Prince of Vladimir" from his Mongol overlords. He cooperated closely with the Mongols and collected tribute from other Russian principalities on their behalf. This relationship enabled Ivan to gain regional ascendancy, particularly over Muscovy's chief rival, the northern city of Tver'. In 1327 the Orthodox metropolitan transferred his residency from Vladimir to Moscow, further enhancing the prestige of the new principality.
In the fourteenth century, the grand princes of Muscovy began gathering Russian lands to increase the population and wealth under their rule (see table 2, Appendix). The most successful practitioner of this process was Ivan III (the Great; r. 1462-1505), who conquered Novgorod in 1478 and Tver' in 1485. Muscovy gained full sovereignty over the ethnically Russian lands in 1480 when Mongol overlordship ended officially, and by the beginning of the sixteenth century virtually all those lands were united. Through inheritance, Ivan obtained part of the province of Ryazan', and the princes of Rostov and Yaroslavl' voluntarily subordinated themselves to him. The northwestern city of Pskov remained independent in this period, but Ivan's son, Vasiliy III (r. 1505-33), later conquered it.
Ivan III was the first Muscovite ruler to use the titles of tsar and "Ruler of all Rus'." Ivan competed with his powerful northwestern rival Lithuania for control over some of the semi-independent former principalities of Kievan Rus' in the upper Dnepr and Donets river basins. Through the defections of some princes, border skirmishes, and a long, inconclusive war with Lithuania that ended only in 1503, Ivan III was able to push westward, and Muscovy tripled in size under his rule.